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LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died. The zoo said Gramma had ongoing bone conditions and that caretakers had to make the difficult ...

The zoo said Gramma had ongoing bone conditions and that caretakers had to make the difficult decision to say goodbye to the legendary resident of the zoo. Gramma was known among her San Diego Zoo ...

She was believed to be around 141 years old when she was euthanized due to "ongoing bone conditions related to advanced age," the zoo said Known as "the Queen of the Zoo," Gramma was in the first ...